Rodríguez Domínguez, U. and Caplan, J. B. (2019). A hexagonal Fourier model of grid cells. Hippocampus, 29, 37–45. [Supplementary materials][Featured cover]


Kato, K. and Caplan, J. B. (in press). Order of items within associations. Journal of Memory and Language.

Madan, C. R., Fujiwara, E., Caplan, J. B. and Sommer, T. (2017). Emotional arousal impairs association-memory: roles of amygdala and hippocampus. NeuroImage, 156, 14–28.

Klingmüller A., Caplan, J. B. and Sommer, T. (in press). Intrusions in episodic memory: reconsolidation or interference? Learning and Memory.

Kato, K. and Caplan, J. B. (in press). The brain’s representations may be compatible with convolution-based memory models. Canadian Journal of Experimental Psychology.

Chen, Y. Y. and Caplan, J. B. (2017). Rhythmic activity and individual variability in recognition memory: theta oscillations correlate with performance whereas alpha oscillations correlate with ERPs. Journal of Cognitive Neuroscience, 29(1), 183–202.

Burton, R. L., Lek, I. and Caplan, J. B. (2017). Associative independence revisited: competition between conflicting associations can be resolved or even reversed in one trial. Quarterly Journal of Experimental Psychology, 70(4), 832-857.


Caplan, J. B. and Madan, C. R. (2016). Word-imageability enhances association-memory by increasing hippocampal engagement. Journal of Cognitive Neuroscience, 28(10), 1522–1538.


Talmi, D., Caplan, J. B., Richards, B. & Moscovitch, M. (2015). Long-term recency in anterograde amnesia. PLoS ONE, 10(6), e0124084.

Caplan, J. B., Bottomley, M., Kang, P., & Dixon, R. A. (2015). Distinguishing rhythmic from non-rhythmic brain activity during rest in healthy neurocognitive aging. NeuroImage, 112, 341-352.

Caplan, J. B. (2015). Order-memory and association-memory. Canadian Journal of Experimental Psychology, 69(3), 221-232.

Caplan, J. B., Madan, C. R., & Bedwell, D. J. (2015). Item properties may influence item-item associations in serial recall. Psychonomic Bulletin & Review, 22(2), 483-489.


Chen, Y. Y., Lithgow, K., Hemmerich, J. A., & Caplan, J. B. (2014). Is what goes in what comes out? Encoding and retrieval event-related potentials together determine memory outcome. Experimental Brain Research, 232(1), 3175-3190.

Liu , Y. S., Chan, M. & Caplan, J. B. (2014). Generality of a congruity effect in judgements of relative order. Memory & Cognition, 42(7), 1086-1105.

Cruikshank, L. C., Caplan, J. B., & Singhal, A. (2014). A perception-based ERP reveals that the magnitude of delay matters for memory-guided reaching. Experimental Brain Research, 232(7), 2087-2094.

Caplan, J. B., Boulton, K. L., & Gagné, C. L. (2014). Associative asymmetry of compound words. Journal of Experimental Psychology: Learning, Memory, & Cognition, 40(4), 1163-1171.

Caplan, J. B., Rehani, M., & Andrews, J. C. (2014). Associations compete directly in memory. Quarterly Journal of Experimental Psychology, 67(5), 955-978.

Masuda, T., Russell, M. J., Chen, Y. Y., Hioki, K., & Caplan, J. B. (2014). N400 incongruity effect in an episodic memory task reveals different strategies for handling relevant contextual information for Japanese than European Canadians. Cognitive Neuroscience, 5(1), 17-25.


Cruikshank, L. C, Caplan, J. B., & Singhal, A. (2012). Human electrophysiological reflections of the recruitment of perceptual processing during actions that engage memory. Journal of Vision, 12(6), 1-13.

Legge, E. L. G., Madan, C. R., Ng, E. T., & Caplan, J. B. (2012). Building a memory palace in minutes: Equivalent memory performance using virtual versus conventional environments with the Method of Loci. Acta Psychologica, 141(3), 380-390.

Madan, C. R., Fujiwara, E., Gerson, B. C., & Caplan, J. B. (2012). High reward makes items easier to remember, but harder to bind to a new temporal context. Frontiers in Integrative Neuroscience, 6, 61.

Madan, C. R., Caplan, J. B., Lau, C. S. M., & Fujiwara, E. (2012). Emotional arousal does not enhance association-memory. Journal of Memory and Language, 66, 695-716.

Cruikshank, L. C., Singhal, A., Hueppelsheuser, M. & Caplan, J. B. (2012). Theta oscillations reflect a putative neural mechanism for human sensorimotor integration. Journal of Neurophysiology, 107(1) 65-77.

Hughes, A. M., Whitten, T. A., Caplan, J. B. & Dickson, C. T. (2012). BOSC: A Better OSCillation detection method, extracts both sustained and transient rhythms from rat hippocampal recordings. Hippocampus, 22(6), 1417-1428.


Rehani, M., & Caplan, J. B. (2011). Interference and the representation of order within associations. Quarterly Journal of Experimental Psychology, 64(7), 1409-1429.

Whitten, T. A., Hughes, A. M., Dickson, C. T., & Caplan, J. B. (2011). A better oscillation detection method robustly extracts EEG rhythms across brain state changes: The human alpha rhythm as a test case. NeuroImage, 54(2), 860-874. [BOSC scripts]


Madan, C. R., Glaholt, M. G., & Caplan, J. B. (2010). The influence of item properties on association-memory. Journal of Memory and Language, 63, 46-63.

Jacobs, J., Korolev, I. O., Caplan, J. B., Ekstrom, A. D., Litt, B., Baltuch, G., Fried, I., Schulze-Bonhage, A., Madsen, J. R., & Kahana, M. J. (2010). Right-lateralized brain oscillations in human spatial navigation. Journal of Cognitive Neuroscience, 22(5), 824-836.


Chan, M., Ross, B., Earle, G., & Caplan, J. B. (2009). Precise instructions determine participants’ memory search strategy in judgments of relative order in short lists. Psychonomics Bulletin & Review, 16(5), 945-951.

Caplan, J. B. , Glaholt, M. G. and McIntosh, A. R. (2009). EEG activity underlying successful study of associative and order information. Journal of Cognitive Neuroscience, 21(7), 1346-1364.


Talmi, D., Anderson, A. K., Riggs, L., Caplan, J. B. and Moscovitch, M. (2008) Immediate memory consequences of the effect of emotion on attention to pictures. Learning & Memory, 15(3), 172-182.


Caplan, J. B. & Glaholt, M. G. (2007). The roles of EEG oscillations in learning relational information. NeuroImage, 38(3), 604-616.

Newman, E. L., Caplan, J. B., Kirschen, M. P., Korolev, I. O., Sekuler, R. and Kahana, M. J. (2007). Learning your way around town: How virtual taxicab drivers learn to use both layout and landmark information. Cognition, 104(2), 231-253.

Caplan, J. B., McIntosh, A. R. and De Rosa, E. (2007) Two distinct functional networks for successful resolution of proactive interference, Cerebral Cortex, 17(7), 1650-1663.


Caplan, J. B., Glaholt, M. G. and McIntosh, A. R. (2006). Linking associative and list memory: pairs versus triples, Journal of Experimental Psychology: Learning, Memory & Cognition, 32(6), 1244-1265.

Caplan, J. B., Luks, T. L., Simpson, G. V., Glaholt, M. and McIntosh, A. R. (2006). Parallel networks operating across attentional deployment and motion processing: A multi-seed partial least squares fMRI study, NeuroImage, 29(4), 1192-1202.


Caplan, J. B. (2005). Associative isolation: unifying associative and order paradigms. The Journal of Mathematical Psychology, 49}(5), 383-402.

Ekstrom, A. D., Caplan, J. B., Shattuck, K., Fried, I. and Kahana, M. J. (2005). Human hippocampal theta activity during virtual navigation, Hippocampus, 15(7), 881-889.


Caplan, J. B. (2004) Unifying models of paired associates and serial learning: insights from simulating a chaining model. NeuroComputing, 58-60, 739-743.


Howard, M. W., Rizzuto, D. S., Caplan, J. B., Madsen, J. R., Lisman J., Aschenbrenner-Scheibe, R., Schulze-Bonhage, A. and Kahana, M. J. (2003) Gamma oscillations correlate with working memory load in humans. Cerebral Cortex, 13}(12), 1369-1374.

Ekstrom, A. D., Kahana, M. J., Caplan, J. B., Fields, T. A., Isham, E. A., Newman, E. L. and Fried, I. (2003) Cellular networks underlying human spatial navigation. Nature, 425(6954), 184-188.

Caplan, J. B., Madsen, J. R., Schulze-Bonhage, A., Aschenbrenner-Scheibe, R., Newman, E. L. and Kahana, M. J. (2003) Human theta oscillations related to sensorimotor integration and spatial learning. Journal of Neuroscience, 23(11), 4726-4736.


Kahana, M. J. and Caplan, J. B. (2002) Associative asymmetry in probed recall of serial lists. Memory & Cognition, 30(6), 841-849.


Caplan, J. B., Kahana, M. J., Raghavachari, S. and Madsen, J. R. (2001) Distinct patterns of brain oscillations underlie two basic parameters of human maze learning. Journal of Neurophysiology, 86(1), 368-380.

Raghavachari, S., Kahana, M. J., Rizzuto, D. S., Caplan, J. B., Kirschen, M. P., Bourgeois, B., Madsen, J. R. and Lisman, J. E. (2001) Gating of human theta oscillations by a working memory task. Journal of Neuroscience, 21(9), 3175-3183.


Caplan, J. B., Kahana, M. J., Sekuler, R., Kirschen, M. and Madsen, J. R. (2000) Task dependence of human theta: the case for multiple cognitive functions. NeuroComputing, 32-33, 659-665.


Kahana, M. J., Caplan, J. B., Sekuler, R. and Madsen, J. (1999) Using intracranial recordings to study theta: Response to J. O'Keefe and N. Burgess (1999). trends in Cognitive Science, 3(11), 406-407.

Kahana, M. J., Sekuler, R, Caplan, J. B., Kirschen, M. and Madsen, J. (1999) Human theta oscillations exhibit task dependence during virtual maze navigation. Nature, 399(6738), 781-784.